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It's simple, the physiology of the trout is different from the warm water fishes and they are in an environment (moving water, susceptibility to temperature changes, existing in areas that are not native habitat) that taxes their systems to begin with, any added stress is over load . Trout played for a long period of time(relative to size of the fish) build up an acid in their system. The longer they are played the more dangerous the acid level of build up. If one fishes in such a manner to force extended playing of the fish they place the trout in greater danger of mortality. In order to properly release such fish, regardless of size, there is an extended period the angler should spend in reviving it. I've to often seen an angler play a fish for an extended period, hold it pointed upstream let it slip away, only to see it come floating by belly up, a short time later. This is a short explanation and some of the variables. I'm sure those with a scientific explanation will be more complete. Respectfully rvreclus

PS. A great read on this subject is "Trout Streams" by Paul R. Needham, a may be a bit dated but a good intro to the subject.